| Crossing over | Linkage |
|---|---|
| $(1)$ It leads to the separation of linked genes. | $(1)$ It keeps the genes together. |
| $(2)$ It involves the exchange of segments between non-sister chromatids of homologous chromosomes. | $(2)$ It involves genes located on the same chromosome. |
| $(3)$ The frequency of crossing over can never exceed $50\%$. | $(3)$ The number of linkage groups can never be more than the haploid chromosome number. |
| $(4)$ It increases variability by forming new gene combinations. | $(4)$ It reduces variability. |
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